When at rest, Homoptera fold the wings roof-wise over
the body, whereas Heteroptera fold them flat and slightly overlapping. The
legs of most bugs are unremarkable, except in a few predatory forms which
have raptorial front legs for catching prey, rather like those of
Mantids, and in some aquatic forms where the hind legs are somewhat
flattened and fringed with hairs for swimming. Cerci are absent,
although many Homoptera (e.g. most aphids, family Aphididae) have a
pair of tube-like structures at the rear end, called cornicles
or siphunculi, from which they can exude a waxy, defensive fluid
to repel predators. Also, in some aquatic Heteroptera (e.g. water
scorpions and their relatives, family Nepidae) there is a long
appendage at the tip of the abdomen, which forms a respiratory siphon
or breathing tube. Metamorphosis is simple, with a variable number
of nymphal stages depending on species. The true bugs form a large Order
with about 70,000 species, of which over 1,500 occur in the British Isles.
Many species are of considerable economic importance as pests, not only
by causing direct damage or injury to plants and animals, but also by
transmitting many viral diseases.
Agricultural and Horticultural Crop Pests
- these include a wide range of homopteran bugs, such as many of the
whiteflies (Aleyrodoidea), the aphids and their allies (Aphidoidea),
the mealybugs and scale insects (Coccoidea), the jumping plant lice
(Psylloidea), and some of the froghoppers or spittlebugs, treehoppers
and leafhoppers (Cicadoidea); as well as several heteropteran species,
particularly among the squash bugs (Coreidae), the capsid bugs
(Miridae), and the shield bugs or stink-bugs (Pentatomidae).
Biting and Blood-sucking Pests - these
include the well known bedbugs (Cimicidae), and a few species of assassin
bugs (Reduviidae) and flower bugs (Anthocoridae).
However, the vast majority of Hemiptera are quite harmless
insects and some of the predatory forms can be regarded as beneficial when they
habitually prey on insect pests. Furthermore, a few species of scale
insects (Coccoidea) are of great value in the production of useful
commodities, such as shellac, cochineal and various waxes, although
nowadays some of these products have been largely replaced by synthetic
materials.
Shellac - a resinous material used in the
manufacture of varnishes, polishes and many other products - is prepared
from a secretion of the Indian Lac-insect, Laccifer lacca. Similarly,
a related insect from Madagascar, Gascardia sp., yields an
inferior type of lac containing some waxes, known as 'gum-lac'.
Cochineal - a red dye used for colouring
foodstuffs, medicines and cosmetics - is extracted from the female Mexican
Cactus Scale, Dactylopius coccus (= Coccus cacti). The insect
lives on one of the prickly-pear cacti of Southern and Central
America, but has been cultivated elsewhere in Southern Europe and
North Africa.
Kermes (also known as granum tinctum)
- a crimson dye, similar to cochineal - comes from the female scales
of Kermes ilicis (= Coccus ilicis), which lives on
the twigs and branches of evergreen oaks in Southern Europe and
North Africa.
Waxes - used locally in China, India and
elsewhere for making candles and medicinal preparations - are obtained
from several Oriental species of homopteran bugs, such as
Ericerus pe-la and Ceroplastes ceriferus.
For those with historical interests, the 'manna' of
biblical stories is believed, by some, to have been the syrupy honeydew
exuded by scale insects and plant lice (in particular the species
Trabutina mannipara, which feeds on the tamarix tree of
Palestine). In warm, dry climates this exudate quickly solidifies
into sugary lumps as the water evaporates, and it is easily collected
as it rains down from trees that are heavily infested by these
insects - perhaps the origin of the saying, 'manna from heaven'. |